Cite as: »Volstorf, Jenny, and Vighnesh Samel. 2026. Set nets (Dossier). In: fair-fish database, ed. fair-fish international association. World Wide Web electronic publication. Version A | 1.0. CC BY 4.0. https://fair-fish-database.net/db/methods/catch/set-nets/dossier/«
General remarks
Set nets are an important fishing gear used globally. Fishers anchor set nets to the sea bed to erect a wall of netting. If set nets consist of a single layer of netting, they are called “gillnets”; set nets consisting of three layers of netting are referred to as “trammel nets”. Due to the thin material the net is almost invisible to the IND, catching them when they swim into it. Fishers let the net soak for some time which can amount to several hours, potentially stressing the IND and subjecting them to predation and scavenging. This can be ameliorated by reducing soak times and using different types of predator deterrent devices. Fishers then haul the caught IND back to the vessel.
Fishers disentangle the caught IND manually, after which they can drop or throw them into storage units – a practice that can lead to stress and injuries. To prevent excessive exposure to air and gravity, it is crucial to stun the IND immediately after disentanglement, followed by rapid slaughter whilst the IND are still unconscious. To the best of our knowledge, this is not in practice at the moment.
To prevent undersized, injured, or surplus target IND from undergoing all hazards of catching, emersion, and disentangling before being discarded, fishers should use mesh sizes that assure good size selectivity given the body ∅ in season and region. And fisheries management can impose restrictions on the fishing depth, net length, catch area, or catch season.
Set nets are not usually species specific. They lead to the bycatch of various non-target taxa. One group – marine megafauna such as big sharks, seabirds, and marine mammals – causes trepidations over its conservation status. Discarding another group – aquatic animals of low commercial value – can be reduced through appropriate mesh sizes, landing obligations, and initiatives to encourage the consumption of these species. Another important issue is the risk of ghost fishing due to abandoned, lost, and discarded nets, which can be addressed by using fewer or biodegradable nets, making reporting of lost gear mandatory, and undertaking gear retrieval attempts.
1 Commercial relevance
Global catch rate: 2,790.7 t/year 2019
Catch in the global ocean (for gillnets): 2,790.7 t/year 2019 or 2.6% 1.
Weight per catch: ~0.013-32 kg/km
Observations Merluccius merluccius: 28 mm mesh size: 0.017 g/m (0.017 kg/km), 30 mm mesh size: 0.25 g/m (0.25 kg/km), 32 mm mesh size: 0.015 g/m (0.015 kg/km) 2, 53 mm mesh size: 5.3 kg/1,300 m (~4 kg/km), 62.5 mm mesh size: 7.4 kg/1,300 m (5.6 kg/km), 80 mm mesh size: 7.7 kg/1,300 m (~5.9 kg/km), 70 mm mesh size: 8.2 kg/1,300 m (~6.3 kg/km) 3, 81 mm mesh size: 95.6 kg/10,000 m (~9.56 kg/km) 4, 90 mm mesh size: 8.9 ± 7.9 kg/750 m (~11.87 kg/km), 80 mm mesh size: 14.3 ± 8.2 kg/750 m (~19 kg/km), 70 mm mesh size: 24 ± 15.4 kg/750 m (~32 kg/km) 5, 28 mm, 30 mm, and 32 mm mesh size: 11.6 kg/600 m (~19.4 kg/km) 6, 40 mm mesh size: 0.27 kg/set, 60 mm mesh size: 1.15 kg/set, 70 mm mesh size: 1.7 kg/set, 80 mm mesh size: 1.22 kg/set, 90 mm mesh size: 0.65 kg/set 7.
Observations Mullus surmuletus: B-type nets: 0.013 kg/km, S-type nets: 5.3 kg/km 8, 62.5 mm mesh size: 0.6 kg/1,300 m (~0.46 kg/km), 53 mm mesh size: 1.5 kg/1,300 m (~1.15 kg/km) 3, for 3 m net height: 87 mm mesh size: 0.17 kg/100 m (~1.70 kg/km), 75 mm mesh size: 0.69 kg/100 m (~6.9 kg/km), 53 mm mesh size: 1.3 kg/100 m (13 kg/km), 60 mm mesh size: 1.6 kg/100 m (16 kg/km), for 5 m net height: 87 mm mesh size: 0.12 kg/100 m (~1.2 kg/km), 75 mm mesh size: 0.55 kg/100 m (~5.5 kg/km), 60 mm mesh size: 1.07 kg/100 m (~10.7 kg/km), 53 mm mesh size: 1.1 kg/100 m (11 kg/km) 9, in mesh 40 mm out mesh 240 mm: 0.15 kg/trial, in mesh 40 mm with selvedge out mesh 240 mm: 0.17 kg/trial, in mesh 36 mm out mesh 240 mm: 0.2 kg/trial, in mesh 36 mm with selvedge out mesh 240 mm: 2.8 kg/trial 10, 36 mm: 0.65 kg/survey 11.
2 Target species
Target species: Merlucciidae, Mullidae (further research needed)
Procedure: a long horizontal wall of netting set at the bottom to catch FISHES swimming into it by snagging, gilling, wedging, or other methods of entanglement
Catch at or near the bottom of the sea with variable soaking times:
Gear: gillnets (single wall of netting) of 107-5,000 m length × 1.5-6.3 m height with mesh sizes 17-150 mm; trammel nets (one net with smaller mesh sizes sandwiched between two outer nets) of 183-5,000 m length × 0.8-1.7 m height with inner mesh sizes: 22-50+ mm, outer mesh sizes: 120-240 mm
Gillnet dimensions (single wall of netting): 107-5,000 m length × 1.5-6.3 m height, trammel net dimensions (one net with smaller mesh sizes sandwiched between two outer nets): 183-5,000 m length × 0.8-1.7 m height:
Observations Merluccius merluccius: gillnets: 107 m length × 5.5 m height 19, 200 m length × 3.4 m height 6, 750 m length × 6.3 m height 5, 1,300 m length × 3.3 m height 18, 1,300 m length × 4 m height 3, 1,600 m length × 5 m height 16, 3,000 m length × ~5.4 m height 4, 5,000 m length × 3 m height 7.
Observations Mullus surmuletus: gillnets: ~183 m length × 2.4 m height 1424, 300 m length × 1.8 m height 11, ~493-914 m length 12, 1,000 m length × 3-5 m height 9, 4,600-5,000 m length × 1.5-3 m height 30; trammel nets: ~183 m length × 0.8-1.7 m height 31, 500 m length 10, B-type nets: 823 m length × 1.6 m height, S-type nets: 919 m length × 1.6 m height 8, <2,000 m length per fisher or <5,000 m length per vessel 32.
Mesh sizes for gillnets (single wall of netting): 17-150 mm; mesh sizes for trammel nets (one net with smaller mesh sizes sandwiched between two outer nets): inner mesh sizes: 22-50+ mm, outer mesh sizes: 120-240 mm:
Observations Merluccius merluccius: gillnets: 26 mm nylon monofilament, hanging ratio of 0.5 18, 53 mm, 62.5 mm, 70 mm, and 82 mm monofilament, hanging ratio of 0.5 3, 28 mm, 30 mm, and 32 mm 2, 40 mm, 60 mm, 70 mm, and 80 mm monofilament polyamide, hanging ratio of 0.5 7, 52-80 mm 33, 70 mm, 80 mm, 90 mm, 100 mm, hanging ratio of ~0.5 5, 81 mm 17, 80 mm, 100 mm, 120 mm, 140 mm monofilament nylon, hanging ratio of 0.6 19, 100-150 mm 16.
Observations Mullus surmuletus: gillnets: 17 mm, 19 mm, 21 mm, and 23 mm, hanging ratio of 0.6 12, 36 mm nylon 11, 53 mm, 60 mm, 75 mm, and 87 mm 9, 50-59 mm, 60-69 mm, and >90 mm 34, 60-100 mm 30, 108 mm 1424; trammel nets: in mesh 22 mm or 28 mm, out mesh 140 mm 13, in mesh 26 mm, out mesh 120 mm 31, B-type nets: in mesh 32-72 mm, S-type nets: in mesh 27 mm 8, in mesh 36-40 mm, out mesh 240 mm 10, in mesh 40-52 mm 33, in mesh >50 mm 32.
Vessels: 4.8-42 m in length belonging to small-scale and industrial fleets
Observations Merluccius merluccius: 7-10.6 m long 30, 10 m long, 85 HP engine 18, 17.5 m long, 195 kW engine 19, ~20 m long 216, 27 m long stern trawler 7.
Observations Mullus surmuletus: 4.8-12.6 m long, 13.5-230 HP engine 8, 6 m long, 9 HP engine 10, 6-12 m long 32, 8 m long, 43 HP engine 11, 9.5 m long, 68 HP engine 9, 12.7 m long, two engines of 74 HP each 12, 30.3-42 m long 35.
4 Prospection
Prospection methods: fish finder device (further research needed)
Mullus surmuletus: fishers use fish finder devices to determine the depth 30, which helps determine the presence of IND36.
4.2 Prospection hazards and mitigation measures
No data found yet.
5 Setting
Setting methods: shore netting (further research needed)
Mullus surmuletus: with shore netting (Oct-Apr), fishers deploy net in dropping tide in bay and trap IND when they want to leave the bay just at a foot of water 1424.
Setting hazards and mitigation measures: probably no hazards (further research needed)
Mullus surmuletus: given that setting does not include active chasing or passive ways to direct IND (noise, light, FADs, etc.), hazard consequences during setting are unlikely 0.
6 Catching/hauling (in the water)
Catching depth range: range 2-700 m
Observations: Merluccius merluccius: 24-250 m, mostly <100 m 7, 38-327 m 16, 50-70 m 26, 65-78 m 5, 68-405 m 18, 95-256 m 3, 150-300 m 5, 500-700 m 4, Mullus surmuletus: 2-68 m 37, 10-65 m 8, 12-70 m 9, 15-21 m 10, 16-86 m 30, 18-60 m 12, 20-80 m 13, 15-30 fathoms (~27.4-54.9 m) 31.
Catching depth hazards and mitigation measures: probable hazards as a combined effect of depth and hauling speed – further research needed on mitigation measures
Merluccius merluccius: catching depth and unknown speed of hauling in nets results in barotrauma 2738282939. Further research needed on how to avoid this.
Mullus surmuletus: given catching depth and unknown speed of hauling, barotrauma is possible 0.
6.3 Hauling speed
No data found yet.
6.4 Hauling speed hazards and mitigation measures
No data found yet. Catching duration: soak time range 1-24+ h with additional variable time of setting and hauling
Observations Merluccius merluccius: soak time of 1.5-3 h (end of lowering to end of picking up: 4.9-6.3 h), sometimes >12 h (end of lowering to end of picking up: 15-16.3 h) 18, 3-24+ h, mean 13-18 h 7, 5 h 3, 7-12 h 4, 10 h 28, average 17.2-22.1 h 16, or 20-22 h 26.
Observations Mullus surmuletus: soak time 60-75 min and hauling of 150 min 30, soaking ~1.5 h 11, soaking 2 h from just before dusk 10, average 172 min soaking from before dawn, average 42 min hauling at sunrise 8, soaking 3-5 h from before sunrise 9, soaking from 2 h prior to 2 h after sunset 12, soaking from sunset to sunrise 31, average 5.8-9.4 h 40.
Catching duration hazards and mitigation measures: long soak time – prefer shorter soak time (further research needed), from Merluccius merluccius: prefer 7 h soak time over 12 h
Merluccius merluccius: in gillnets in 500-700 m depth, condition of IND decreased with increasing soak time: from 77% IND in good condition after 7 h soak time to 35% in good condition after 12 h, probably due to scavenging crustaceans 4. Further research needed on types of hazard consequences.
Mullus surmuletus: with bottom-set gillnets, after soak time of 60-75 min and hauling of 150 min, almost 50% IND arrived on board inactive, another ca 25% were lethargic with maximum duration of activity 4 min 30. During soaking, interactions with 1-12 bottlenose dolphins Tursiops truncatus 12/88 times (13.6%), during hauling 17/88 times (19.3%), spending average 20 min and reducing catch of M. surmuletus. Negative correlation between catch volume and duration of soaking time and interactions with dolphins 8.
Catching gear contact hazards and mitigation measures: snagging, gilling, wedging, or entangling – not avoidable; inappropriate mesh sizes resulting in catching undersized IND (or for Merluccius merluccius: large sexually productive females) – prefer mesh size that assures good size selectivity given body ∅ in season and region
Snagging, gilling, wedging, or entangling – not avoidable:
Merluccius merluccius: given the principle of set nets to catch IND2526272829 by the head region (called "snagging") 5, by the gills (called "gilling") 51936, by the mid body (called “wedging”) 536, or by wrapping the whole body or attaching protruding body parts like teeth (called “entangling”) 5193, hazard consequences are probable 0. Further research needed on types of hazard consequences.
Mullus surmuletus: given the principle of set nets to catch IND2021222314 by the head region (called “snagging”) 41, by the gills (called "gilling") 14, by the mid body (called “wedging”) 41, or by wrapping the whole body or attaching protruding body parts like teeth (called “entangling”) 41, hazard consequences are probable 0. During catching, substantial scale loss (26-50% of body surface) in IND, indicating a high risk of external injuries 30. Catching SPAWNERS with gillnets and transportation caused skin damage resulting in mortality within 2-3 weeks; more IND dead without external injuries 42.
Inappropriate mesh sizes resulting in catching undersized IND (or for Merluccius merluccius: large sexually productive females) – prefer mesh size that assures good size selectivity given body ∅ in season and region:
Merluccius merluccius: the following mesh sizes assure good size selectivity, i.e., the listed sizes of IND will have contact with the gear: gillnets in 500-700 m depth after 7-12 h soak time: 81 mm mesh size (commercially used) caught IND of 30-65 cm TOTAL LENGTH and avoided catch of IND <27 cm minimum landing size, avoiding bycatch 4. Commercial gillnets in 150-300 m depth: decreasing number of IND and increasing catching length with increasing mesh size (69.5, 81, 88.5 mm), no difference between 88.5 mm commercial and 96.3 mm experimental mesh size. Decreasing number of males, increasing number of females with increasing length. Of 17-65 cm TOTAL LENGTH caught, increasing frequency of IND <27 cm minimal catch size with increasing mesh size (0.6-2.1%), but also lower number of IND <45.3 cm size at maturity 43↶5, prompting recommendation to increase minimum mesh size to 81 mm even though minimum mesh size cannot avoid entangling of small IND by teeth 5. Experimental gillnets in mostly <100 m depth: highest number and weight of IND at mesh size 60, 70, 80 mm with best combination at 70 mm; largest IND but lower number at 90 mm. Of 19-67 cm TOTAL LENGTH caught, 5% IND <27 cm (minimal landing size) at 60-80 mm mesh size 7. Corresponds with legally enforced mesh size ≥80 mm given maturity in females at 45.3 cm 43↶7. In experimental bottom-set gillnets after average 23.3 h soak time, tendency of increasing size of caught IND with increasing mesh size (80, 100, 120 mm) when caught by gills (majority of cases); no effect when caught by teeth. Lower catch rate at 140 mm mesh size. At 120 mm mesh size (commercial size), small frequency of IND <60 cm and majority of IND 70-85 cm, indicating good size selectivity of currently used mesh size 19. Experimental gillnets in 95-256 m depth: tendency of increasing weight and length and decreasing number of IND with increasing mesh size (53, 62.5, 70, 82 mm). Best combination and lowest risk for catching large sexually productive females at 62.5 mm – larger than the commercially used 53 mm mesh size 3. Bottom-set gillnets with 28, 30, 32 mm mesh size in 50-70 m depth after 22 h soak time: highest weight and number of IND caught and of 23.5-41.0 cm TOTAL LENGTH, tendency of lowest percentage (2.7%) of IND <25 cm minimal landing size 44↶2 at 30 mm mesh size (commercially used), indicating good size selectivity of currently used mesh size 2. With bottom gillnets in 50-70 m depth at soak time 20-22 h, no difference in catching size between mesh sizes 28, 30, 33 mm: 23.5-41 cm TOTAL LENGTH, indicating that commercial mesh size 30 mm assures exceeding minimal landing size 6 of 20 cm TOTAL LENGTH in Turkey 45↶6. The following mesh sizes potentially cause bycatch, i.e., they should be avoided to prevent those IND from having contact with the gear: 70 mm commercial mesh size with highest number of IND caught, but also high number of IND <45.3 cm size at maturity 43↶5 and high discard and scavenging rate in gillnets in 150-300 m depth, prompting recommendation to cease using 70 mm mesh size commercially 5. Of 19-67 cm TOTAL LENGTH caught, IND <27 cm (minimal landing size) mainly at 40 mm mesh size in experimental gillnets in mostly <100 m depth, decreasing percentage with increasing mesh size 7. At 80 mm mesh size, IND <60 cm caught in experimental bottom-set gillnets after average 23.3 h soak time, increasing size of caught IND with increasing mesh size 19. No IND <20 cm minimal landing size and no males below size at maturity (19 cm TOTAL LENGTH46↶3), but 32.5% immature females (<35.1 cm TOTAL LENGTH46↶3) caught at 53 mm mesh size (commercially used). Higher number of larger (potentially very sexually productive) females than males at mesh sizes 70 and 82 mm mean potentially big risk to overexploitation of population in experimental gillnets in 95-256 m depth 3. Lowest number of IND and weight caught and tendency of highest percentage (4.4%) of IND <25 cm minimal landing size 44↶2 at 32 mm mesh size in bottom-set gillnets in 50-70 m depth after 22 h soak time 2. Assuming maturity in females at 21.5 cm, in males at 25.7-27.7 cm 47↶648↶6, females were caught above length at maturity, but males were caught below length at maturity in 4.5-8% at mesh sizes 28, 30, 32 mm with bottom gillnets in 50-70 m depth at soak time 20-22 h 6.
Mullus surmuletus: the following mesh sizes assure good size selectivity, i.e., the listed sizes of IND will have contact with the gear: bottom-set gillnets at 18-60 m depth, after soaking from 2 h prior to 2 h after sunset: decreasing number of IND and increasing catching length with increasing mesh size (17, 19, 21, 23 mm). Of 10-21 cm FORK LENGTH caught, all IND were >10 cm FORK LENGTH minimal landing size 49↶12, indicating good size selectivity of currently used mesh sizes and better size selectivity than bottom trawls 12. Gillnets at 20 m isobath with mesh size 36 mm, after soaking for 1.5 h, caught IND of 9.4-23.1 cm TOTAL LENGTH, mostly above minimum conservation reference size (11 cm 50↶11) and partly below 50% size at maturity (15.5 cm 50↶11) 11. Recommended mesh size for Mediterranean (38.5 mm 50↶11) might eliminate these issues 0. Gillnets at 12-70 m, after soaking 3-5 h from before sunrise: decreasing number of IND with increasing mesh size (53, 60, 75, 87 mm); highest weight at 60 mm. Of 18-39 cm TOTAL LENGTH caught, all IND were >15 cm minimal landing size, prompting recommendation to increase allowed mesh size from 50 to 60 mm. No difference in number of IND and weight with 5 m high net instead which is not allowed and cannot be recommended 9. Trammel nets at 15-21 m depth with 36-40 mm in mesh and 240 mm out mesh, after soaking for 2 h caught IND of 14.1-21.7 cm TOTAL LENGTH, all >11 cm minimal landing size 10. Trammel nets with 40-52 mm mesh size caught IND of 11-27 cm TOTAL LENGTH, mostly above size at maturity (15-16 cm 51↶33) and with better size selectivity than bottom trawls (5-23 cm TOTAL LENGTH) 33. The following mesh sizes potentially cause bycatch, i.e., they should be avoided to prevent those IND from having contact with the gear: With bottom-set gillnets, mesh size 50-90+ mm, during 2009-2015: <5% bycatch of IND >40 g (ca 18 cm TOTAL LENGTH) minimal commercial landing weight, for trammel nets >15%, probably indicating a risk for overexploitation of the population 34. Trammel nets with minimum mesh size 50 mm, during 2000-2012 caught IND of 9-29 cm TOTAL LENGTH, partly <14 cm size at 50% first maturity, probably indicating a risk for overexploitation of the population 32.
Other catching gear hazards and mitigation measures: probably no crowding
Merluccius merluccius: no crowding step before hauling the net on board, but IND remained caught and with the same distance to other con-specifics under and above water when net was continuously hauled 2526272829, probably not resulting in decreasing distance to neighbour 0.
Mullus surmuletus: no crowding step before hauling the net on board, but IND remained caught and with the same distance to other con-specifics under and above water when net was continuously hauled 20212223 or when IND were collected lying on the shore 1424, probably not resulting in decreasing distance to neighbour 0.
Environmental parameter changes during catching: abrupt salinity change possible (further research needed)
Merluccius merluccius: in ca 25 m depth, low salinity (ca 18 ppt) water from Black Sea meets high salinity (ca 38 ppt) water from Mediterranean 6.
Parameter change hazards and mitigation measures: abrupt salinity change possible – further research needed on mitigation measures
Merluccius merluccius: by fishing depth of 50-70 m 6, IND are hauled from low salinity to high salinity layer, and hazard consequences are probable 0. Further research needed on the rate M. merluccius adapts to osmotic changes without decreasing welfare and on reports of osmoregulatory distress after gillnetting.
Predation pressure: probably scavengers and predators during soaking
Merluccius merluccius: probably scavenging crustaceans during soaking 4. Depredation rate by Tursiops truncatus 25%/haul 40.
Mullus surmuletus: probably scavenging crustaceans 14 or depradation by cetaceans 11 during soaking. Depredation rate by Tursiops truncatus ~21-25%/haul 40. Interactions with Tursiops truncatus reduced catch of M. surmuletus8. Bite injuries 8, probably by cuttlefishes Sepia spp., common octopus Octopus vulgaris, European conger Conger conger, Mediterranean moray Murena helena53↶8.
Predation hazards and mitigation measures: probably scavengers and predators during soaking – prefer shorter soak time, from Merluccius merluccius: prefer 7 h soak time over 12 h (further research needed), from Mullus surmuletus: amend or switch gear to avoid recognisable sounds, close fishing for certain times or areas or switch up areas, apply "stealth fishing", attach dolphin deterrent devices or, favourably, dolphin interactive devices to the gear
Merluccius merluccius: during soaking, depredation rate by Tursiops truncatus 25%/haul 40. Given that scavenged IND are discarded after manual disentangling 26, scavenging should be avoided or at least decreased 0. In gillnets in 500-700 m depth, condition of IND decreased with increasing soak time: from 77% IND in good condition after 7 h soak time to 35% in good condition after 12 h, probably due to scavenging crustaceans 4. Further research needed on types of hazard consequences.
Mullus surmuletus: during soaking, IND may be damaged 2111, probably by scavenging crustaceans 14 or predating cetaceans 11. Depredation rate by Tursiops truncatus ~21-25%/haul 40. Given that scavenged IND are most likely discarded after manual disentangling, scavenging should be avoided or at least decreased 0. During soaking, interactions with 1-12 bottlenose dolphins Tursiops truncatus 12/88 times (13.6%), during hauling 17/88 times (19.3%), spending average 20 min and reducing catch of M. surmuletus. Negative correlation between catch volume and duration of soak time and interactions with dolphins 8 indicates probable reduction of lost catch through lower soak time 0. Of those IND still caught in the net when hauled on board, without interaction with dolphins, mainly bite injuries 8, probably by cuttlefishes Sepia spp, common octopus Octopus vulgaris, European conger Conger conger, Mediterranean moray Murena helena53↶8; with interaction with dolphins, roughly equal amount of bite injuries, only head remaining, only tail remaining, and fragments remaining 8. As dolphins are probably lured by sounds that the gear make, recommended to amend or switch up gear; dolphins could also hear if vessels leave for the sea at night, so recommended to close fishing for certain times or areas or switch up areas 8 or apply "stealth fishing" 54↶8. Using dolphin deterrent devices reduced depradation by T. truncatus by 48%, using dolphin interactive devices reduced depradation by 50%, but their efficiency gradually reduces over time 55.
7 Emersion (transfer from water)
Emersion/transfer methods: hauling the net, collecting from the shore
Emersion gear contact hazards and mitigation measures: emerge snagged, gilled, wedged, or entangled – not avoidable; gear contact during emersion – further research needed on mitigation measures
Emerge snagged, gilled, wedged, or entangled:
Merluccius merluccius: given the principle of set nets to catch IND2526272829 by the head region (called "snagging") 5, by the gills (called "gilling") 51936, by the mid body (called “wedging”) 536, or by wrapping the whole body or attaching protruding body parts like teeth (called “entangling”) 5193, hazard consequences are probable 0. Further research needed on types of hazard consequences.
Mullus surmuletus: given the principle of set nets to catch IND2021222314 by the head region (called “snagging”) 41, by the gills (called "gilling") 14, by the mid body (called “wedging”) 41, or by wrapping the whole body or attaching protruding body parts like teeth (called “entangling”) 41, hazard consequences are probable 0. During catching, substantial scale loss (26-50% of body surface) in IND, indicating a high risk of external injuries 30. Catching SPAWNERS with gillnets and transportation caused skin damage resulting in mortality within 2-3 weeks; more IND dead without external injuries 42.
Gear contact during emersion – further research needed on mitigation measures:
Merluccius merluccius: when the net is hauled on board, IND may be catapulted on deck, and given how the effect of gravity interacts with the type of catching (snagging, gilling, wedging, entangling) and therefore how the net affects the skin and given contact with several reels along which the net is guided 2526272829, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: when the net is hauled onboard, IND may be catapulted on deck, and given how the effect of gravity interacts with the type of catching (snagging, gilling, wedging, entangling) and therefore how the net affects the skin and given contact with several reels along which the net is guided 20212223 and assuming IND are not already dead, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them. Catching SPAWNERS with gillnets and transportation caused skin damage resulting in mortality within 2-3 weeks; more IND dead without external injuries 42.
Other emersion gear hazards and mitigation measures: probably no crowding
Merluccius merluccius: no crowding step before hauling the net on board, but IND remained caught and with the same distance to other con-specifics under and above water when net was continuously hauled 2526272829, probably not resulting in decreasing distance to neighbour 0.
Mullus surmuletus: no crowding step before hauling the net on board, but IND remained caught and with the same distance to other con-specifics under and above water when net was continuously hauled 20212223 or when IND were collected lying on the shore 1424, probably not resulting in decreasing distance to neighbour 0.
Predation pressure: probably low (further research needed)
Mullus surmuletus: predation pressure during emersion is probably low 0.
Predation hazards and mitigation measures: probably low predation during emersion (rather during soaking; further research needed)
Merluccius merluccius: given short distance between water surface and aboard vessel and thus hardly any occasion 2526272829, predation pressure during emersion is probably low 0. More time and more convenient occasion during soak time or hauling → D2.
Mullus surmuletus: given short distance between water surface and aboard vessel and thus hardly any occasion 202223, predation pressure during emersion is probably low 0. More time and more convenient occasion during soak time or hauling → D2.
Environmental parameter changes during emersion: exposure to air and gravity
Mullus surmuletus: IND are exposed to gravity when the net is hauled on board 20212223 or when they are collected lying on the shore 1424.
Parameter change hazards and mitigation measures: exposure to air and gravity – further research needed on mitigation measures
Air exposure – further research needed on mitigation measures:
Merluccius merluccius: given that when the net is hauled on board, IND are exposed to air 2526272829, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: given that when the net is hauled on board 20212223 or the IND are collected lying on the shore 1424, IND are exposed to air and assuming IND are not already dead, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid them.
Gravity exposure/experience own weight – further research needed on mitigation measures:
Merluccius merluccius: given that when the net is hauled on board 2526272829, IND are exposed to gravity and given no evolutionary adaptation to experiencing own weight in air 56, hazard consequences following emersion are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: given that when the net is hauled on board 20212223 or the IND are collected lying on the shore 1424, IND are exposed to gravity and given no evolutionary adaptation to experiencing own weight in air 56, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Release hazards and mitigation measures: rough handling, dropping, air exposure – further research needed on mitigation measures; to prevent handling for undersized IND, prefer mesh size that avoids catching undersized IND given body ∅ in season and region
Rough handling – further research needed on mitigation measures; to prevent handling for undersized IND, prefer mesh size that avoids catching undersized IND given body ∅ in season and region (→ D1):
Merluccius merluccius: given that manual disentangling happens fast and rough 2526272829, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them. Given that undersized IND are discarded after manual disentangling 252729, hazard consequences are probable 0. Further research needed on types of hazard consequences.
Mullus surmuletus: given that manual disentangling happens fast and rough 2021222330 and assuming IND are not already dead, hazard consequences are probable 300. Further research needed on types of hazard consequences and on how to avoid or decrease them. Given that undersized IND are discarded after manual disentangling 23 and assuming IND are not already dead, hazard consequences are probable 0. Further research needed on types of hazard consequences.
Dropping – further research needed on mitigation measures:
Merluccius merluccius: given that during manual disentangling, IND may fall to the floor 29 and that after disentangling, IND are thrown into trays 252729, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: given that during manual disentangling, IND may fall to the floor 202223 and that after disentangling, IND are thrown into buckets 202122301424 and assuming IND are not already dead, hazard consequences are probable 300. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Air exposure – further research needed on mitigation measures:
Merluccius merluccius: given that during manual disentangling, IND are exposed to air 2526272829, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: given that during manual disentangling, IND are exposed to air 20212223301424 and assuming IND are not already dead, hazard consequences are probable 300. Further research needed on types of hazard consequences and on how to avoid or decrease them.
9 Sorting
Sorting methods: manual sorting during disentangling (further research needed):
Sorting hazards and mitigation measures: rough handling during disentangling, dropping, air exposure – further research needed on mitigation measures; to prevent handling for undersized IND, prefer mesh size that avoids catching undersized IND given body ∅ in season and region
Rough handling – further research needed on mitigation measures; to prevent handling for undersized IND, prefer mesh size that avoids catching undersized IND given body ∅ in season and region (→ D1):
Merluccius merluccius: given that manual disentangling happens fast and rough 2526272829, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them. Given that undersized IND are discarded after manual disentangling 252729, hazard consequences are probable 0. Further research needed on types of hazard consequences.
Mullus surmuletus: given that manual disentangling happens fast and rough 2021222330 and assuming IND are not already dead, hazard consequences are probable 300. Further research needed on types of hazard consequences and on how to avoid or decrease them. Given that undersized IND are discarded after manual disentangling 23 and assuming IND are not already dead, hazard consequences are probable 0. Further research needed on types of hazard consequences.
Dropping – further research needed on mitigation measures:
Merluccius merluccius: given that during manual disentangling, IND may fall to the floor 29 and that after disentangling, IND are thrown into trays 252729, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: given that during manual disentangling, IND may fall to the floor 202223 and that after disentangling, IND are thrown into buckets 202122301424 and assuming IND are not already dead, hazard consequences are probable 300. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Air exposure – further research needed on mitigation measures:
Merluccius merluccius: given that during manual disentangling, IND are exposed to air 2526272829, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: given that during manual disentangling, IND are exposed to air 20212223301424 and assuming IND are not already dead, hazard consequences are probable 300. Further research needed on types of hazard consequences and on how to avoid or decrease them.
10 Storing
Storing methods: live storage, storage after death (further research needed)
Merluccius merluccius: IND were immediately gutted alive and placed in cleaning bath after disentangling and sorting, then transferred to storing containers initially without ice 252627, eventually with ice 28.
Storing gear contact hazards and mitigation measures: contact with holding container – prefer immediate stunning followed by slaughter while still unconscious (further research needed)
Merluccius merluccius: IND were immediately gutted alive and placed in cleaning bath after disentangling and sorting 252627. Given that in busy times with a full net, gutting might take place only after a waiting time in a holding container, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Mullus surmuletus: given that IND were thrown into buckets 202122301424 with water 22, later placed on and under ice 1424, and assuming IND are not already dead, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Other storing gear hazards and mitigation measures: exposure to air/lack of oxygen and ice, crowding – prefer immediate stunning followed by slaughter while still unconscious (further research needed)
Merluccius merluccius: IND were immediately gutted alive and placed in cleaning bath after disentangling and sorting 252627. Given that in busy times with a full net, gutting might take place only after a waiting time in a holding container, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them. The IND were then transferred to storing containers initially without ice 252627, eventually with ice 28. Given that IND are already dead when stored in containers, hazard consequences are improbable 0.
Mullus surmuletus: given that IND were thrown into buckets 202122301424 with water 22, later placed on and under ice 1424, and assuming IND are not already dead, hazard consequences are probable 0. Further research needed on types of hazard consequences and on how to avoid or decrease them.
11 Stunning and slaughter
Stunning methods: none – further research needed on the efficacy of percussive stunning followed by gill cut or gutting
Merluccius merluccius: given that IND were immediately gutted alive after disentangling and sorting without prior stunning 252627, hazard consequences are probable 0. Further research needed on types of hazard consequences and on quick and painless stunning methods.
Mullus surmuletus: given that IND were thrown into buckets 202122301424 with water 22, later placed on and under ice 1424, probably resulting in asphyxia 300 or hypothermia 0. Recommended to percussively stun with a fishing priest or captive bolt pistol, followed by cutting gill, immersing in ice water, or storing on ice 57.
Slaughter methods: asphyxia and/or hypothermia – from Merluccius merluccius: gutting
Merluccius merluccius: IND were immediately gutted alive after disentangling and sorting without prior stunning 252627 and placed in ice 28, which prevents live storage and prolonged suffering. Given that stunning is lacking, hazard consequences are probable 0. Further research needed on types of hazard consequences and on quick and painless stunning methods.
Mullus surmuletus: given that IND were thrown into buckets 202122301424 with water 22, later placed on and under ice 1424, probably resulting in asphyxia 300 or hypothermia 0. Recommended to percussively stun with a fishing priest or captive bolt pistol, followed by cutting gill, immersing in ice water, or storing on ice 57.
12 Bycatch/discard
12.1 Bycatch/discard rate
Target species: from low (Mullus surmuletus: 1.4%) to such high discard rate (Merluccius merluccius: 42%) that gillnets were banned (further research needed)
Merluccius merluccius: gillnets in 500-700 m depth: 42% biomass of caught IND were discarded due to being partially or completely deteriorated by scavenging crustaceans. Condition decreased with increasing soak time: from 77% IND in good condition after 7 h soak time to 35% in good condition after 12 h 4. Higher discard rate from gillnetting than longline (42% versus 7% biomass) in 500-700 m depth, together with lower catch and yield and lower quality, led to ban of gillnets in favour of longlines 4.
Mullus surmuletus: discarding probability is assumed to be low due to high commercial value of M. surmuletus58↶5960↶59. Gillnets at 12-70 m, after soaking 3-5 h from before sunrise: at commercial mesh size 53 mm, 1.4% discarded due to damage 9.
Non-target species: 0.2-84% of non-usable bycatch, 2.4-72.5% of potentially usable bycatch (commercially relevant species), <10.9% fed the crew, <53.3% used as bait
Merluccius merluccius: given the principle of set nets to be passive gear floating or standing in habitat that other species use or traverse, there is a risk to also catch co-existing, preyed-on, and predating species 0.
Usable bycatch: bottom gillnets in northern Atlantic off south-west of England and Ireland in 38-327 m depth with 100-150 mm, mostly 120 mm mesh sizes, caught Pollachius pollachius, Pollachius virens, Molva molva, Gadus morhua besides target M. merluccius16. Gillnets off the southern Portuguese coast (the Algarve) in 500-700 m depth with 81 mm mesh size (commercially used): 14% of biomass commercially valuable of which very commonly Micromesistius poutassou, commonly Galeus melastomus, uncommonly Brama brama, Phycis blennoides, Ruvettus pretiosus, Scyliorhinus canicula, rarely Auxis rochei, Beryx decadactylus, Helicolenus dactylopterus, Lepidopus caudatus, Lophius piscatorius, Raja clavata, Nephrops norvegicus4. Multispecies fisheries off western Portuguese coast, experimental gillnets in mostly <100 m depth: commercial species Pagellus acarne, Mullus surmuletus, Trisopterus luscus, Trachurus trachurus, almost no bycatch at 90 mm 7. In experimental gillnets in northern Tyrrhenian sea in 95-256 m depth with mesh sizes 53, 62.5, 70, 82 mm: decreasing biomass of Trachurus trachurus (72.5% → 36.1%), increasing biomass of Chelidonichthys lucerna (2.4-9.8%), small biomass of 16-20 other species 3. Bottom-set gillnets in south-eastern Sea of Marmara in 50-70 m depth with mesh size 28, 30, 32 mm contained commercial species Solea solea (39.4%) 2.
Non-usable bycatch: bottom gillnets in northern Atlantic off south-west of England and Ireland in 38-327 m depth with 100-150 mm, mostly 120 mm mesh sizes, caught Phocoena phocoena (0.4-0.7 IND/trip or 15 IND/1,000 km net set), which were already dead or dying, detangled themselves or got detangled, and Delphinus delphis. Per year, estimated 2,200 Phocoena phocoena in vessels ≥15 m or 6.2% of porpoise population in Celtic sea 16 exceeds proposed 1% rate 61↶16. Gillnets off the southern Portuguese coast (the Algarve) in 500-700 m depth and 81 mm mesh size (commercially used): 1% of biomass non-commercial species of which commonly Todaropsis eblanae, rarely Benthodesmus elongatus, Centrolophus niger, Chimaera monstrosa, Dalatias licha, Etmopterus pusillus, Etmopterus spinax, Hoplostethus mediterraneus, Hymenocephalus italicus, Naucrates ductor, Illex coindetii4. Multispecies fisheries off western Portuguese coast, experimental gillnets in mostly <100 m depth: at mesh size 40 mm mainly (84%) low (Dicologlossa cuneata, Scyliorhinus canicula) or non-commercial species (Citharus linguatula) 7. Bottom-set gillnets in south-eastern Sea of Marmara in 50-70 m depth with mesh size 28, 30, 32 mm contained non-commercial species Raja clavata (0.1%), Buglossidium luteum (0.1%) which were discarded and Trachurus mediterraneus (8.2%), Chelidonichthys lucerna (2.5%), Lophius piscatorius (0.2%) which fed the crew 2. Some bycatch (mackerel) fed the crew 26, large bycatch was disentangled and discarded 2729.
Mullus surmuletus: given the principle of set nets to be passive gear floating or standing in habitat that other species use or traverse, there is a risk to also catch co-existing, preyed-on, and predating species 0.
Usable bycatch: with gillnets of 300 m length × 1.8 m height, 36 mm mesh size, in northern Aegean Sea, Greece: 20.7% biomass catch of (mostly) JUVENILES of Pagellus acarne, Diplodus annularis, and Pagellus erythrinus threatened these species' populations 11. With gillnets of 1,000 m length × 3 m height and commercial mesh size 53 mm, in Asturias, northwest Spain: M. surmuletus caught together with other species of high economic value representing 33.5% IND of catch (mostly Merluccius merluccius, some Serranus cabrilla, Pagellus acarne, rarely Trisopterus luscus, Pagellus erythrinus, Sepia officinalis, Pagellus bogaraveo, Belone belone, Octopus vulgaris, Trachurus trachurus, Auxis rochei, Chelidonichthys lucerna, Coris julis, Oblada melanura, Pollachius pollachius, Sarda sarda, Scorpaena porcus, Symphodus melops, Zeus faber), another 53.3% IND for use as bait (Trachurus trachurus, Boops boops) 9. With gillnets, in Northern Atlantic, at 68 mm mesh size: pollocks, streaked gurnards, cuckoo wrasses, lesser spotted dogfishes 2223, scads, poutings, lings, one octopus, horse mackerels 22, one sea cucumber 23. With shore netting, in Northern Atlantic, at 108 mm mesh size: bycatch of bass, Gilthead seabream (Sparus aurata), mullet 1424, jellyfish 14. With trammel nets of 500 m length and mesh sizes in 36-40 mm, out 240 mm, in Izmir Bay, Aegean Sea, Turkey: bycatch of commercial species (mostly Diplodus annularis, some Boops boops, Spicara flexuosa, Diplodus vulgaris, Pagellus erythrinus, rarely Sepia officinalis, Octopus vulgaris, Trachurus trachurus, Trigla lucerna, Pagellus acarne) 10.
Non-usable bycatch: with gillnets with 60-75 mm mesh size, off the southern Portuguese coast (the Algarve): ~0.9% bycatch of Tursiops truncatus40. With gillnets of 1,000 m length × 3 m height and commercial mesh size 53 mm, in Asturias, northwest Spain: 13.2% IND discarded due to commercial species being damaged (36.3% IND) or being undersized (21.1% IND) or of too little numbers (12.7% IND) or exceeding total allowable catch (4.6% IND) or due to non-commercial species (25.3% IND) (mostly Polybius henslowi, Sardina pilchardus, Boops boops, Trachurus trachurus, some Trachinus draco, Chelidonichthys cululus, Scorpaena scrofa, Trisopterus luscus, rarely Scomber colias, Scorpaena notata, Callionymus lyra, Serranus cabrilla, Chelidonichthys obscurus, Scomber scombrus, Arnoglossus imperialis, Pagellus acarne, Trisopterus minutus, Chelidonichthys gurnadus, Corystes cassivelaunus, Diplodus vulgaris, Echinaster sepositus, Engraulis encrasicolus, Holothuria forskali, Labrus bergylta, Labrus mixtus, Octopus vulgaris, Pagellus bogaraveo, Sarda sarda, Scyliorhynus canicula, Symphodus bailloni) 9. With trammel nets of 500 m length and mesh sizes in 36-40 mm, out 240 mm, in Izmir Bay, Aegean Sea, Turkey: 77.4-81.4% IND discarded (Anomura spp., Maja squinado, Hexaplex trunculus, Serranus scriba, Symphodus tinca, Arnoglossus materna, Coris julis, Bolinus brandaris, Aporrhais pespelecani, Serranus cabrilla, Eledone moschata, Sardina pilchardus, Symphodus ocellanus, Serranus hepatus, Blennius ocellaris, Gobius niger, Chromis chromis) 10.
General bycatch: with bottom-set trammel nets of 500-2,000 m length × 1.6 m height, 27 mm mesh size, off Sardinia: the following other non-target species caught of which unclear whether they are used (commercially or at sea) or discarded: mostly Diplodus vulgaris, Scorpaena porcus, Diplodus annularis, Scorpaena scrofa, Serranus cabrilla, Pagellus erythrinus, Sepia officinalis, some Symphodus tinca, Phycis phycis, Sciena umbra, Spodyliosoma cantharus, Spicara maena, Sepia orbignyana, Symphodus roissali, Dentex dentex, rarely Octopus vulgaris, Labrus merula, Diplodus puntazzo, Trigloporus lastoviza, Uranoscopus scaber, Zeus faber, Boops boops, Labrus viridis, Sarpa salpa, Palinurus elephas, Diplodus sargus, Scyliorhinus canicula, Trachinus draco, Trisopterus minutus capelanus, Loligo vulgaris, Pagellus acarne, Conger conger, Symphodus mediterraneus, Dasyatis pastinaca, Labrus micaculatus, Sepia elegans, Sphyraena sphyraena, Seriola dumerili, Trachinus radiatus, Trachinus araneus, Scomberesox saurus, Raja clavata, Raja brachyura, Maja squinado8.
12.2 Bycatch/discard hazards
Target species: probable hazards as a combined effect of depth and hauling speed; long soak time; snagging, gilling, wedging, or entangling; inappropriate mesh sizes resulting in catching undersized IND (or for Merluccius merluccius: large sexually productive females); abrupt salinity change possible; probably scavengers and predators during soaking; gear contact during emersion; exposure to air and gravity; rough handling, dropping, air exposure; displacement; predation pressure
Probable hazards as a combined effect of depth and hauling speed →D3.
Merluccius merluccius: given that surviving discarded IND will have to return to their natural living depth, vertical displacement is probable 56. Further research needed on types of hazard consequences.
Mullus surmuletus: given that surviving discarded IND will have to return to their natural living depth, vertical displacement is probable 56. Further research needed on types of hazard consequences.
Predation pressure:
Merluccius merluccius: given that surviving discarded IND are probably disoriented, stressed, and weakened after the catching process and release, and given that seabirds and other predators might gather near fishing boats, predation pressure is probable 56. Further research needed on how to avoid or decrease this.
Mullus surmuletus: given that surviving discarded IND are probably disoriented, stressed, and weakened after the catching process and release 56, and given that seabirds 202123 and other predators might gather near fishing boats, predation pressure is probable 56. Predation on several FISHES62, including M. surmuletus63, by seabirds like Morus bassanus and by Larus spp. 62. Further research needed on types of hazard consequences.
Non-target species: risk of overexploitation, injuries/mortality
risk of overexploitation:
Mullus surmuletus: usable bycatch: given that in northern Aegean Sea, Greece, fishing season (spring-autumn) fell in reproductive season for these species, risk for overexploitation of the population due to extraction of SPAWNERS of Diplodus annularis, Mullus barbatus, Sphyraena sphyraena, and Trachurus trachurus is probable 11. With gillnets of 1,000 m length × 3 m height and commercial mesh size 53 mm, in Asturias, northwest Spain: M. surmuletus caught together with other species of high economic value representing 33.5% IND of catch, another 53.3% IND for use as bait (Trachurus trachurus, Boops boops). Higher number of IND and weight of catch used for bait with 5 m high net instead which is not allowed and cannot be recommended 9.
Mullus surmuletus: non-usable bycatch: with gillnets of 1,000 m length × 3 m height and commercial mesh size 53 mm, in Asturias, northwest Spain: 13.2% IND discarded due to commercial species being damaged (36.3% IND) or being undersized (21.1% IND) or of too little numbers (12.7% IND) or exceeding total allowable catch (4.6% IND) or due to non-commercial species (25.3% IND). Higher number of IND damaged and higher number of non-commercial species with 5 m high net instead which is not allowed and cannot be recommended 9. Highest discards in May-Oct, probably due to more FISHES inhabiting seagrass 10.
injuries/mortality:
Merluccius merluccius: non-usable bycatch: some bycatch (mackerel) fed the crew 26, large bycatch was disentangled and discarded 2729, sometimes after being heavily tossed to the floor 26 (probable makeshift stunning 0) or half-eaten by predator 26 which makes survival improbable 0.
Mullus surmuletus: non-usable bycatch: FISHES used as bait (Trachurus trachurus, Boops boops) are not treated carefully during disentangling due to their low appreciation 9.
Target species: probably scavengers and predators during soaking – from Mullus surmuletus: close fishing for certain times or areas or switch up areas; high discard rate – from Merluccius merluccius: potentially prefer longlines over gillnets (further research needed); risk of overexploitation – from Mullus surmuletus: prefer set nets over bottom trawl and possibly gillnets over trammel nets; other management measures: total allowable catch, regulate fishing days per year, closure areas or seasons, from Merluccius merluccius: regulate net length, from Mullus surmuletus: limit fishing depth (<50 m)
Probably scavengers and predators during soaking – from Mullus surmuletus: close fishing for certain times or areas or switch up areas →D6.
High discard rate – from Mullus surmuletus: potentially prefer longlines over gillnets (further research needed):
Merluccius merluccius: higher discard rate from gillnetting than longline (42% versus 7% biomass) in 500-700 m depth, together with lower catch and yield and lower quality, led to ban of gillnets in favour of longlines 4.
Risk of overexploitation – from Mullus surmuletus: prefer set nets over bottom trawl and possibly gillnets over trammel nets:
Mullus surmuletus: with bottom-set gillnets, mesh size 50-90+ mm, off France, during 2009-2015: <5% bycatch of IND >40 g (ca 18 cm TOTAL LENGTH) minimal commercial landing weight; for trammel nets >15%, probably indicating a risk for overexploitation of the population 34 In Mediterranean, bottom trawl was more detrimental for population of M. surmuletus than gillnets 12 or trammel nets 33 due to worse size selectivity 12 including JUVENILES33.
Risk of overexploitation – total allowable catch, regulate fishing days per year, closure areas or seasons, from Merluccius merluccius: regulate net length, from Mullus surmuletus: limit fishing depth (<50 m):
Merluccius merluccius: total allowable catch, closure of areas to prevent catch of JUVENILES because minimum mesh size cannot avoid entangling of small IND by teeth 5. Fishing season off the southern Portuguese coast (the Algarve): first semester 5, in Sea of Marmara: April-May 2, March-May 6. To avoid overexploitation, recommended to regulate length of nets, fishing days per year, close areas or seasons, especially during catching season winter-spring which corresponds with reproductive period in northern Tyrrhenian sea 3. In simulation of management strategies for Mediterranean coast off Spain, only 83% reduction in fishing days for multispecies and multigear fisheries (set nets, longline, bottom trawl) during 5 years of simulation achieved maximum sustainable yield, which the authors consider an unrealistic decrease in fishery activity throughout the year 64.
Mullus surmuletus: in another DEMERSAL species caught with set nets in the Northern Atlantic and Mediterranean, Merluccius merluccius, total allowable catch is recommended and closure of areas to prevent catch of JUVENILES because minimum mesh size cannot avoid entangling of small IND by teeth 5 which M. surmuletus loses in the upper jaw only from >9 cm TOTAL LENGTH on 65. Further research needed to determine whether this applies to M. surmuletus as well. Fishing season: Mar-Aug 33, spring-autumn 11, May-July up to year round 37, July-Dec 32, Sep-Dec 8, Oct-Apr 1424, year round 103431. In M. merluccius, to avoid overexploitation, recommended to regulate fishing days per year, close areas or seasons, especially during the spawning season 3. Further research needed to determine whether this applies to M. surmuletus as well. Trammel nets off Egypt, Mediterranean, caught IND of 9.1-30 cm TOTAL LENGTH which is partly <15.4 cm age at maturity and thus a risk for overexploitation, since the JUVENILES cannot reproduce before being caught. Possible mitigation measures need to be researched: larger mesh size, closure of fishing areas 66. Management regulations for trammel nets, off Mallorca: limitation to 6 fishing days per week, forbidden to catch >50 m depth 32
Non-target species: risk of overexploitation – from Mullus surmuletus: adjust minimum conservation reference size, encourage consumption of non-commercial species, landing obligation; injuries/mortality – from Mullus surmuletus: encourage consumption of non-commercial species
Risk of overexploitation – from Mullus surmuletus: adjust minimum conservation reference size, encourage consumption of non-commercial species, landing obligation:
Mullus surmuletus: usable bycatch: recommended to specify lengths at maturity and adjust minimum conservation reference size 11. Encouraging consumption of species (so far) used as bait and implementing a landing obligation could reduce the amount of bycatch 9.
Injuries/mortality – encourage consumption of non-commercial species:
Mullus surmuletus: non-usable bycatch: implementing a landing obligation and encouraging consumption of (so far) non-commercial species could reduce the amount of discards – also because FISHES used as bait (Trachurus trachurus, Boops boops) are not treated carefully during disentangling due to their low appreciation 9.
12.4 Bycatch avoidance/discard protection in water
Target species: probable hazards as a combined effect of depth and hauling speed – further research needed on mitigation measures; long soak time – prefer shorter soak time (further research needed), from Merluccius merluccius: prefer 7 h soak time over 12 h; snagging, gilling, wedging, or entangling – not avoidable; inappropriate mesh sizes resulting in catching undersized IND (or for Merluccius merluccius: large sexually productive females) – prefer mesh size that assures good size selectivity given body ∅ in season and region; abrupt salinity change possible – further research needed on mitigation measures; probably scavengers and predators during soaking – prefer shorter soak time, from Merluccius merluccius: prefer 7 h soak time over 12 h (further research needed), from Mullus surmuletus: amend or switch gear to avoid recognisable sounds, apply "stealth fishing", attach dolphin deterrent devices or, favourably, dolphin interactive devices to the gear; gear contact during emersion – further research needed on mitigation measures; exposure to air and gravity – further research needed on mitigation measures; rough handling, dropping, air exposure – further research needed on mitigation measures; displacement – further research needed on mitigation measures; predation pressure – from Mullus surmuletus: use visual scarybird device as deterrent against predatory seabirds
Probable hazards as a combined effect of depth and hauling speed – further research needed on mitigation measures →D3
Long soak time – prefer shorter soak time (further research needed), from Merluccius merluccius: prefer 7 h soak time over 12 h →D4.
Snagging, gilling, wedging, or entangling – not avoidable →D1.
Inappropriate mesh sizes resulting in catching undersized IND (or for Merluccius merluccius: large sexually productive females) – prefer mesh size that assures good size selectivity given body ∅ in season and region →D1.
Abrupt salinity change possible – further research needed on mitigation measures →D5.
Probably scavengers and predators during soaking – prefer shorter soak time, from Merluccius merluccius: prefer 7 h soak time over 12 h (further research needed), from Mullus surmuletus: amend or switch gear to avoid recognisable sounds, apply "stealth fishing", attach dolphin deterrent devices or, favourably, dolphin interactive devices to the gear →D6.
Gear contact during emersion – further research needed on mitigation measures →D7.
Exposure to air and gravity – further research needed on mitigation measures →D8.
Rough handling, dropping, air exposure – further research needed on mitigation measures →D9.
Displacement – further research needed on mitigation measures: Merluccius merluccius: given that surviving discarded IND will have to return to their natural living depth, vertical displacement is probable 56. Further research needed on types of hazard consequences and on how to avoid or decrease them.
Predation pressure – from Mullus surmuletus: use visual scarybird device as deterrent against predatory seabirds:
Merluccius merluccius: given that surviving discarded IND are probably disoriented, stressed, and weakened after the catching process and release, and given that seabirds and other predators might gather near fishing boats, predation pressure is probable 56. Further research needed on how to avoid or decrease this.
Mullus surmuletus: given that surviving discarded IND are probably disoriented, stressed, and weakened after the catching process and release 56, and given that seabirds 202123 and other predators might gather near fishing boats, predation pressure is probable 56. Predation on several FISHES62, including M. surmuletus63, by seabirds like Morus bassanus – which was reduced through use of a visual deterrent in the form of a bird of prey ("scarybird") – and by Larus spp. – which was reduced through retaining to-be-discarded IND in buckets until after hauling the net 62. Given the use of small buckets resulting in crowded conditions, hazard consequences are probable 0. Further research needed on types of hazard consequences.
Non-target species: risk of overexploitation – from Mullus surmuletus: install guarding net (selvedge) for trammel nets
Merluccius merluccius: non-usable bycatch: hazard consequences including mortality are best avoided by preventing bycatch 0. Further research needed on gear settings and on other co-existing, preyed-on, and predating species.
Mullus surmuletus: non-usable bycatch: highest discards in May-Oct, probably due to more FISHES inhabiting seagrass. Attaching guarding nets (selvedge) decreased discards to 18.6-22.6%, especially of decapod crustaceans Anomoura spp. and Maja squinado and gastropods Hexaplex trunculus, Bolinus brandaris, Aporrhais pespelecani10.
13 Environmental hazards
13.1 Live bait
No data found yet. Habitat hazards and mitigation measures: seabed damage or impact on benthos relatively low (further research needed)
Merluccius merluccius: catching takes place at the bottom 16192640. The net is usually not dragged along the seafloor 67, though, so seabed damage or huge impact on benthos is relatively low 0.
Mullus surmuletus: given that M. surmuletus is a BENTHIC carnivore 68, catching takes place at the bottom 311283440146213553024. The net is not dragged along the seafloor 1424, though, so seabed damage or huge impact on benthos is relatively low 0. Recommended to monitor the benthic habitat before deploying the net 36.
Ghost fishing risk, gear loss: 2,963 km2/year
Estimate of gillnets abandoned, lost, or discarded globally: average 2,963 km2/year, range 1,153.1-4,772.7 km2/year, average 3,153 m2 gillnets/vessel, corresponding to 0.8% of gillnets 69.
Ghost fishing hazards and mitigation measures: ghost fishing – implement a Code of Good Practice, close areas where active and passive gear interact for one gear at a time, use biodegradable nets, reduce number of nets, soak time, and vessels, make reporting of lost gear mandatory, regularly attempt gear recovery, apply gear retrieval during DEMERSAL trawl surveys
Merluccius merluccius: when gillnets get dragged by a trawl, they may get lost and become "ghost nets". Deliberately "lost" gillnets off the southern Portuguese coast (the Algarve) in 65-78 m depth with mesh size 81 mm (commercially used) started getting holes after 3 months, torn lines after 75 days when set in September or after 248 days when set in spring. May be dragged along the bottom. Catch of number of M. merlucciusIND decreased by 5% after 30 days compared to normally fishing gillnets with 24 h soak time, 35% after 90 days, probably due to biofouling on net, amount of hake remains, or strong currents. No more catches after 248 days when set in May, after 75 days when set in autumn. Per year, estimated 7,139 IND or 2,437 t M. merluccius ghost fished off Algarve 17. Prominent decrease in number of IND after 30 days, gradual decrease until after 230 days, residual bycatch after 12 months when study stopped. 18 species of which most frequently by number Citharus linguatula (34.2-38.7%), Scorpaena notata (23.7%) in spring, Scomber scombrus (30.7%) in autumn, most frequently by weight Lophius piscatorius, Microchirus azevia, Scorpaena notata (in sum 58.1%) in spring, Scomber scombrus, Auxis rochei, Scyliorhinus canicula (in sum 68.1%) in autumn. After 8 months, almost exclusively BENTHIC species. Per year, estimated 20,367 IND or 5.2 t ghost fished off the southern Portuguese coast (the Algarve), equalling 17,213 Euro. Estimated ghost fishing impact of ~25 FISHES or 64.4 kg/100 m of net in ~430 days of active fishing life of the ghost net. To reduce ghost fishing and the loss of nets, the following measures are recommended: implementing a Code of Good Practice, closure of areas where active and passive gear interact for one gear at a time, biodegradable nets, lower number of nets, soak time, and vessels, making reporting of lost gear mandatory, regular gear recovery attempts, gear retrieval during DEMERSAL trawl surveys 70.
Mullus surmuletus: in ghost-net experiments (gillnet: 60 mm mesh size, trammel net: in mesh 60 mm and out mesh 100) off the southern Portuguese coast (the Algarve), that catch several FISHES including M. surmuletus, 15-20 weeks effective fishing lifetime of the net. The nets caught 39 species belonging to 4 main groups (crustaceans: Galathea strigosa, Maja squinado, Macropipus puber, Scyllarus arctus; gastropods: Charonia lampas; molluscs: Cybiun olla, Sepia officinalis; FISHES: Alistes carolinensis, Parablennius gattorugine, Callionymus lyra, Caranx rhonchus, Trachurus trachurus, Conger conger, Phycis phycis, Trisopterus luscus, Pomadasys incisus, Labrus merula, Symphodus bailloni, Symphodus spp., Merluccius merluccius, Mugil spp., Mullus surmuletus, Scomber spp., Scomber japonicus, Scorpaena notata, Serranus spp., Serranus cabrilla, Buglossidium luteum, Dentex gibbosus, Diplodus annularis, Diplodus bellottii, Diplodus cervinus, Diplodus puntazzo, Diplodus sargus, Diplodus spp., Diplodus vulgaris). Estimated 344 FISHES/100 m length caught in 120 days. Probable scavenging and predation by Conger spp., Muraeinadae, and Coris julis on the ghost netted FISHES, especially soft bodied species like M. sumuletus71.
Glossary
BENTHIC = living at the bottom of a body of water, able to rest on the floor DEMERSAL = living and feeding on or near the bottom of a body of water, mostly benthopelagic, some benthic FAD = fish-aggregating device. Artificial or natural floating or anchored device meant to attract small fishes to take shelter and large fishes to prey on them so that fishers have them conveniently aggregated for fishing FISHES = using "fishes" instead of "fish" for more than one individual - whether of the same species or not - is inspired by Jonathan Balcombe who proposed this usage in his book "What a fish knows". By referring to a group as "fishes", we acknowledge the individuals with their personalities and needs instead of an anonymous mass of "fish". FORK LENGTH = from snout to fork of caudal fin as compared to total length (from snout to tip of caudal fin) 52 or standard length (from head to base of tail fin) or body length (from the base of the eye notch to the posterior end of the telson) HP = horse power IND = individuals JUVENILES = fully developed but immature individuals SPAWNERS = adults during the spawning season; in farms: adults that are kept as broodstock TOTAL LENGTH = from snout to tip of caudal fin as compared to fork length (from snout to fork of caudal fin) 52 or standard length (from head to base of tail fin) or body length (from the base of the eye notch to the posterior end of the telson)
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By Jefferson Moser - Moser, Jefferson F. (1902) Salmon Investigations of the Steamer Albatross in the Summer of 1900, Bulletin of the United States Fish Commission, vol.21, 1901, Washington, D.C.: Government Printing Office, Public Domain, https://commons.wikimedia.org/w/index.php?curid=42840920
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Global Fishing Watch. 2026. Apparent fishing effort by set gillnets, worldwide, 2012-2026. Accessed on 02.03.2026. https://globalfishingwatch.org
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Probably, we updated the profile. Check the version number in the head of the page. For more information on the version, see the FAQ about this. Why do we update profiles? Not just do we want to include new research that has come out, but we are continuously developing the database itself. For example, we changed the structure of entries in criteria or we added explanations for scores in the WelfareCheck | farm. And we are always refining our scoring rules.
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The centre of the Overview is an array of criteria covering basic features and behaviours of the species. Each of this information comes from our literature search on the species. If we researched a full Dossier on the species, probably all criteria in the Overview will be covered and thus filled. This was our way to go when we first set up the database.
Because Dossiers are time consuming to research, we switched to focusing on WelfareChecks. These are much shorter profiles covering just 10 criteria we deemed important when it comes to behaviour and welfare in aquaculture (and lately fisheries, too). Also, WelfareChecks contain the assessment of the welfare potential of a species which has become the main feature of the fair-fish database over time. Because WelfareChecks do not cover as many criteria as a Dossier, we don't have the information to fill all blanks in the Overview, as this information is "not investigated by us yet".
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See the question "What does "not investigated by us yet" mean?". In short, if we have not had a look in the literature - or in other words, if we have not investigated a criterion - we cannot know the data. If we have already checked the literature on a criterion and could not find anything, it is "no data found yet". You spotted a "no data found yet" where you know data exists? Get in touch with us at ffdb@fair-fish.net!
Behind an entry, what does "[x]↶[y]" mean?
Once you have clicked on "show details", the entry for a criterion will unfold and display the summarised information we collected from the scientific literature – complete with the reference(s).
As reference style we chose "Springer Humanities (numeric, brackets)" which presents itself in the database as a number in a grey box. Mouse over the box to see the reference; click on it to jump to the bibliography at the bottom of the page. But what does "[x]↶[y]" refer to?
This is the way we mark secondary citations. In this case, we read reference "y", but not reference "x", and cite "x" as mentioned in "y". We try to avoid citing secondary references as best as possible and instead read the original source ourselves. Sometimes we have to resort to citing secondarily, though, when the original source is: a) very old or not (digitally) available for other reasons, b) in a language no one in the team understands. Seldomly, it also happens that we are running out of time on a profile and cannot afford to read the original. As mentioned, though, we try to avoid it, as citing mistakes may always happen (and we don't want to copy the mistake) and as misunderstandings may occur by interpreting the secondarily cited information incorrectly.
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It is also possible, though, that we did have to make a decision between several papers on the same topic. If there are too many papers on one issue than we manage to read in time, we have to select a sample. On certain topics that currently attract a lot of attention, it might be beneficial to opt for the more recent papers; on other topics, especially in basic research on behaviour in the wild, the older papers might be the go-to source.
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This number, for example "C | 2.1 (2022-11-02)", contains 4 parts:
"C" marks the appearance – the design level – of the profile part. In WelfareChecks | farm, appearance "C" is our most recent one with consistent age class and label (WILD, FARM, LAB) structure across all criteria.
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".1" marks the number of minor updates within this appearance. Here, it is minor update 1. With minor updates, we mean changes in formatting, grammar, orthography. It can also mean adding new papers, but if these papers only confirm the score and don't change it, it will be "minor" in our book. With a change to a new appearance, the minor update will be re-set to 0.
"(2022-11-02)" is the date of the last change – be it the initial release of the part, a minor, or a major update. The nature of the changes you may find out in the changelog next to the version number.
If an Advice, for example, has an initial release date and then just a minor update date due to link corrections, it means that – apart from correcting links – the Advice has not been updated in a major way since its initial release. Please take this into account when consulting any part of the database.
General
How to use the FAQ?
First up, you will find answers to questions for the specific page you are on. Scrolling down in the FAQ window, there are also answers to more general questions. Explore our website and the other sub pages and find there the answers to questions relevant for those pages.
How do I navigate in the fair-fish database?
In the fair-fish database, when you have chosen a species (either by searching in the search bar or in the species tree), the landing page is an Overview, introducing the most important information to know about the species that we have come across during our literatures search, including common names, images, distribution, habitat and growth characteristics, swimming aspects, reproduction, social behaviour but also handling details. To dive deeper, visit the Dossier where we collect all available ethological findings (and more) on the most important aspects during the life course, both biologically and concerning the habitat. In contrast to the Overview, we present the findings in more detail citing the scientific references.
Depending on whether the species is farmed or wild caught, you will be interested in different branches of the database.
Farm branch
Founded in 2013, the farm branch of the fair-fish database focuses on farmed aquatic species.
Catch branch
Founded in 2022, the catch branch of the fair-fish database focuses on wild-caught aquatic species.
What is the farm branch?
The heart of the farm branch of the fair-fish database is the welfare assessment – or WelfareCheck | farm – resulting in the WelfareScore | farm for each species. The WelfareCheck | farm is a condensed assessment of the species' likelihood and potential for good welfare in aquaculture, based on welfare-related findings for 10 crucial criteria (home range, depth range, migration, reproduction, aggregation, aggression, substrate, stress, malformations, slaughter).
For those species with a Dossier, we conclude to-be-preferred farming conditions in the Advice | farm. They are not meant to be as detailed as a rearing manual but instead, challenge current farming standards and often take the form of what not to do.
What is the catch branch?
In parallel to farm, the main element of the catch branch of the fair-fish database is the welfare assessment – or WelfareCheck | catch – with the WelfareScore | catch for each species caught with a specific catching method. The WelfareCheck | catch, too, is a condensed assessment of the species' likelihood and potential for good welfare – or better yet avoidance of decrease of good welfare – this time in fisheries. We base this on findings on welfare hazards in 10 steps along the catching process (prospection, setting, catching, emersion, release from gear, bycatch avoidance, sorting, discarding, storing, slaughter).
In contrast to the farm profiles, in the catch branch we assess the welfare separately for each method that the focus species is caught with. In the case of a species exclusively caught with one method, there will be one WelfareCheck, whereas in other species, there will be as many WelfareChecks as there are methods to catch the species with.
Summarising our findings of all WelfareChecks | catch for one species in Advice | catch, we conclude which catching method is the least welfare threatening for this species and which changes to the gear or the catching process will potentially result in improvements of welfare.
What do we mean by "welfare"?
Welfare of aquatic species is at the heart of the fair-fish database. In our definition of welfare, we follow Broom (1986): “The welfare of an individual is its state as regards its attempts to cope with its environment.” Thus, welfare may be perceived as a continuum on which an individual rates “good” or “poor” or everything in between.
We pursue what could be called a combination of not only a) valuing the freedom from injuries and stress (function-based approach) but b) supporting attempts to provide rewarding experiences and cognitive challenges (feelings-based approach) as well as c) arguing for enclosures that mimic the wild habitat as best as possible and allow for natural behaviour (nature-based approach).
What if I have a question?
Try mousing over the element you are interested in - oftentimes you will find explanations this way. If not, there will be FAQ on many of the sub-pages with answers to questions that apply to the respective sub-page. If your question is not among those, contact us at ffdb@fair-fish.net.
Where is the FishEthoBase?
It's right here! We decided to re-name it to fair-fish database for several reasons. The database has grown beyond dealing purely with ethology, more towards welfare in general – and so much more. Also, the partners fair-fish and FishEthoGroup decided to re-organise their partnership. While maintaining our friendship, we also desire for greater independence. So, the name "fair-fish database" establishes it as a fair-fish endeavour.