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Adriatic sturgeon

Acipenser naccarii

Acipenser naccarii (Adriatic sturgeon)
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Distribution
Distribution
Distribution map: Acipenser naccarii (Adriatic sturgeon)

critically endangered



Information


Authors: Paolo Panizzon, João L. Saraiva

Version: C | 1.0
Published: 2026-09-28

Please note: This part of the profile is currently being revised.


Reviewer: Jenny Volstorf
Editor: Jenny Volstorf

Version information:
  • Initial release: 2017-10-27
  • Appearance version: 2026-09-28
  • Major version: 2026-09-28

Cite as: »Panizzon, Paolo, and João L. Saraiva. 2026. Acipenser naccarii (WelfareCheck | farm). In: fair-fish database, ed. fair-fish international association. World Wide Web electronic publication. Version C | 1.0. CC BY 4.0. https://fair-fish-database.net/db/species/acipenser-naccarii/farm/welfarecheck/«





WelfareScore | farm

Acipenser naccarii
LiPoCe
Criteria


Legend

The score card gives our welfare assessments for aquatic species in 10 criteria.

For each criterion, we score the probability to experience good welfare under minimal farming conditions ("Likelihood") and under high-standard farming conditions ("Potential") representing the worst and best case scenario. The third dimension scores how certain we are of our assessments based on the number and quality of sources we found ("Certainty").

The WelfareScore sums just the "High" scores in each dimension. Although good welfare ("High") seems not possible in some criteria, there could be at least a potential improvement from low to medium welfare (indicated by ➚ and the number of criteria).

  • Li = Likelihood that the individuals of the species experience good welfare under minimal farming conditions
  • Po = Potential of the individuals of the species to experience good welfare under high-standard farming conditions
    ➚ = potential improvements not reaching "High"
  • Ce = Certainty of our findings in Likelihood and Potential

WelfareScore = Sum of criteria scoring "High" (max. 10 per dimension)

score-legend
High
score-legend
Medium
score-legend
Low
score-legend
Unclear
score-legend
No findings



General remarks

Acipenser naccarii (Adriatic sturgeon) is a member of the family Acipenseridae. It is a large, long-lived species. Historically, its range included all the rivers of the Adriatic Sea basin, with some possible but unconfirmed populations in some Spanish rivers. Wild populations have declined strongly, mainly due to overfishing and the disturbance of its natural environment. There is uncertainty related to wild ecology, as its presence is nowadays confirmed only in a few, small populations in the Po river basin.

Adriatic sturgeon is currently farmed in Spain and possibly Italy, mainly for the eggs (caviar), secondarily for the meat, depending on each country’s internal market. For this reason, its aquaculture focuses on rearing individuals for a long period of time (6-7 years) in big tanks or earthen ponds until sexual maturity is reached and it is possible to collect the eggs. To this end, hormonal manipulation is used to maximise egg production; individuals destined for meat production are slaughtered before sexual maturation begins.

The WelfareScore is generally low to unclear, reflecting both welfare challenges and substantial gaps in the available evidence. The species’ space needs cannot be fully assessed, as information is mainly available from farms, and further research on home range and depth use in the wild is needed. The presence of a reported resident population in the Ticino river is promising for the Potential of migratory needs, but more research is needed. Reproduction in captivity depends on hormonal manipulation and stripping, while information on breeding behaviour in the wild is lacking. Information on natural density and species-specific aggression is also scarce, although aggression might be limited to the fry stage, as reported in related species. Earthen ponds can provide more natural bottom conditions for juveniles and adults, although natural reproduction and spawning substrate are not currently achievable for spawners. Stress associated with water quality, handling, and density is likely to occur under minimal farming conditions, while mitigation approaches for high-standard farming conditions still require verification in the farming context. Malformations may occur, but evidence on their frequency across life stages in farms is insufficient. Slaughter practices range from spiking without prior stunning to percussive stunning followed by bleeding while the fish remains unconscious, with the latter providing a substantially better welfare outcome when correctly applied. Overall, the limited amount of evidence, particularly on the species’ natural ecology and behaviour, creates considerable uncertainty in several criteria. Further research could substantially improve the assessment and identify more opportunities for welfare improvement.




1  Home range

Many species traverse in a limited horizontal space (even if just for a certain period of time per year); the home range may be described as a species' understanding of its environment (i.e., its cognitive map) for the most important resources it needs access to.

What is the probability of providing the species' whole home range in captivity?

It is unclear for minimal and high-standard farming conditions, given we exclusively found data from farms. Our conclusion is based on a low amount of evidence, as further research is needed on home range in the wild.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS: does not apply.

LARVAE and FRY:

  • WILD: no data found yet.
  • FARM: circular tanks: 0.4 m2 (0.7 m ∅) (restocking facility) 1, fibreglass tanks: 1 m2 2, polyester tanks: 2.2 m2 (5.5 × 0.4 m) 3. Raceways: 0.8-1.4 m2 (0.4 × 2-3.6 m) (restocking facility) 1. Trays or troughs: 2-4 m2 4.
  • LAB: does not apply.

JUVENILES:

  • WILD: no data found yet.
  • FARM: tanks: 7.4 m3 5, fibreglass tanks: 4 m2 2 6, indoor tanks: 800 m2 (100 × 8 m) (restocking facility) 1. Raceways or earthen ponds: 125-1,050 m2 (25-70 × 5-15 m) 4.
  • LAB: does not apply.

ADULTS:

  • WILD: no data found yet.
  • FARM: outdoor concrete tanks: 50 m2 5. Raceways or earthen ponds: 125-1,050 m2 (25-70 × 5-15 m) 4.
  • LAB: does not apply.

SPAWNERS:

  • WILD: no data found yet.
  • FARM: indoor tanks: 800 m2 (100 × 8 m) (restocking facility) 1.
  • LAB: does not apply.



2  Depth range

Given the availability of resources (food, shelter) or the need to avoid predators, species spend their time within a certain depth range.

What is the probability of providing the species' whole depth range in captivity?

It is low for minimal farming conditions, as tanks, ponds, and raceways do not cover the higher end of the depth range in the wild, although we cannot be sure in some age classes. It is unclear for high-standard farming conditions, as these systems do not overlap with the depth range in the wild for JUVENILES and ADULTS but are conceivable to overlap with the depth range in the wild for SPAWNERS. Our conclusion is based on a low amount of evidence, as further research is needed on specific depth information in the wild.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS:

  • WILD: 2-10 m 7↶1.
  • FARM: no data found yet.
  • LAB: does not apply.

LARVAE and FRY:

  • WILD: no data found yet.
  • FARM: circular tanks: 0.1 m (restocking facility) 1, polyester tanks: 0.1 m 3, fibreglass tanks: 0.3 m 2.
  • LAB: does not apply.

JUVENILES:

  • WILD: BENTHIC 8. River pools: found at 10-40 m 9.
  • FARM: fibreglass tanks: 0.3 m 2. Raceways or earthen ponds: <2 m 4.
  • LAB: does not apply.

ADULTS:

  • WILD: → JUVENILES.
  • FARM: raceways or earthen ponds: <2 m 4.
  • LAB: does not apply.

SPAWNERS:

  • WILD: spawning on the riverbed at 2-10 m 7↶1.
  • FARM: raceways or earthen ponds: <2 m 4.
  • LAB: does not apply.



3  Migration

Some species undergo seasonal changes of environments for different purposes (feeding, spawning, etc.), and to move there, they migrate for more or less extensive distances.

What is the probability of providing farming conditions that are compatible with the migrating or habitat-changing behaviour of the species?

It is low for minimal farming conditions, as the species – probably especially the ANADROMOUS, SEMI-ANADROMOUS, and POTAMODROMOUS strains - undertakes more or less extensive migrations, and we cannot be sure that providing each age class with their respective environmental conditions will satisfy their urge to migrate or whether they need to experience the transition. It is medium for high-standard farming conditions, as the space range of the resident strain in captivity potentially covers the migration distance (although hard to compare given different units in WILD and FARM) and as being resident indicates a general adaptability to limited space in tanks, ponds, and raceways. Our conclusion is based on a medium amount of evidence, as further research is needed.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

Facultatively ANADROMOUS or SEMI-ANADROMOUS 1. Some populations POTAMODROMOUS or resident 10 1.

EGGS: does not apply.

LARVAE and FRY:

  • WILD: fresh water 9.
  • FARM: for details of holding systems → F1 and F2.
  • LAB: no data found yet.

JUVENILES:

  • WILD: in the Ticino River, released IND moved average 8.7 km over 16 years, range <1-25.6 km 10. In the Po River, released IND moved average 40.3 km over 16 years, maximum at 254.8 km 10. Present in estuaries 8 9, occasionally in coastal sea waters 9.
  • FARM: for details of holding systems → F1 and F2.
  • LAB: no data found yet.

ADULTS:

  • WILD: →JUVENILES.
  • FARM: for details of holding systems → F1 and F2.
  • LAB: no data found yet.

SPAWNERS:

  • WILD: spawn in rivers 9. Spawning migration March-October 1.
  • FARM: for details of holding systems → F1 and F2.
  • LAB: no data found yet.



4  Reproduction

A species reproduces at a certain age, season, and sex ratio and possibly involving courtship rituals.

What is the probability of the species reproducing naturally in captivity without manipulation of these circumstances?

It is low for minimal and high-standard farming conditions, as the species is manipulated (hormonal manipulation, stripping), and even if stripping is done under anaesthesia, it remains far from natural spawning. Our conclusion is based on a low amount of evidence, as further research is needed on mating type, sex ratio, and courtship rituals in the wild as well as number of spawning events and types of manipulation in captivity.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS: does not apply.

LARVAE and FRY: does not apply.

JUVENILES: does not apply.

ADULTS: does not apply.

SPAWNERS:

  • WILD: males mature at 6-8 8 or 7-11 years old 1, females mature at 12-14 years old 8 1. Spawn April-June 1 11. Females can spawn every 2-4 years 12↶1. For spawning migration → F3, for spawning substrate → F4.
  • FARM: hormonal manipulation to induce spawning 13↶14 15 9 16. Modern techniques avoid slaughter to extract the EGGS, probably including ultrasound for sexing and assessing maturity and manual stripping 13↶14 17, all under anaesthesia 17. Farmers avoid natural reproduction because of impracticability of collecting the EGGS as IND spread them widely 16.
  • LAB: no data found yet.



5  Aggregation

Species differ in the way they co-exist with conspecifics or other species - from being solitary to aggregating in unstructured groups, casually roaming in shoals, or closely coordinating in schools of varying densities.

What is the probability of providing farming conditions that are compatible with the aggregation behaviour of the species?

It is unclear for minimal and high-standard farming conditions, given we mainly found data from farms. Our conclusion is based on a low amount of evidence, as further research is needed on specific density information in the wild and for SPAWNERS in farms.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS: does not apply.

LARVAE and FRY:

  • WILD: no data found yet.
  • FARM: no data found yet.
  • LAB: no data found yet.

JUVENILES:

  • WILD: for Acipenser baerii, at the end of summer, IND in some populations aggregate in wintering pits at the bottom of rivers 18. Further research needed to determine whether this applies to Acipenser naccarii as well.
  • FARM: for sturgeons in Western Europe, tanks: 30-40 kg/m2 19; ponds: 1-5 kg/m2 19. Further research needed to determine whether this applies to Acipenser naccarii as well. High-standard farms: <10 kg/m3 4. Low densities might reduce stress 16.
  • LAB: difficult assessment of stress due to large ranges of cortisol levels 20; no increase in blood cortisol or glucose in IND kept at 17 °C when subjected to stress (crowding for 24 h at 16 kg/m3, long handling; sampled after anaesthesia) 20.

ADULTS:

  • WILD: → JUVENILES.
  • FARM: low densities might reduce stress 16.
  • LAB: no data found yet.

SPAWNERS:

  • WILD: no data found yet.
  • FARM: low densities might reduce stress 16.
  • LAB: no data found yet.



6  Aggression

There is a range of adverse reactions in species, spanning from being relatively indifferent towards others to defending valuable resources (e.g., food, territory, mates) to actively attacking opponents.

What is the probability of the species being non-aggressive and non-territorial in captivity?

It is unclear for minimal farming conditions, given we just found hints on aggression – even cannibalism – in FRY of related species. It is high for high-standard farming conditions, as there is no aggression reported from JUVENILES on (whose welfare we prioritise for scoring over FRY, as JUVENILES-SPAWNERS spend longer time in captivity than FRY). Our conclusion is based on a low amount of evidence, as further research is needed specifically addressing aggression (or lack thereof) as well as species-specific mitigators, should FRY be found to be aggressive.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS: does not apply.

LARVAE and FRY:

  • WILD: no data found yet.
  • FARM: for Acipenser baerii, cannibalistic 21 at 9-15 DPH 22, but low density and correct grading can solve aggression issues 4; for Acipenser gueldenstaedtii, size grading reduced cannibalism rates from 9.3% to 2-5.1% 23. Further research needed to determine wheteher this applies to Acipenser naccarii as well.
  • LAB: no data found yet.

JUVENILES:

  • WILD: no data found yet.
  • FARM: not aggressive under high-standard farming conditions 4 16.
  • LAB: for Acipenser fulvescens housed together after 1 week of acclimatisation, no aggressive interactions with each other over 12 hours 24. Further research needed to determine whether this applies to Acipenser naccarii as well.

ADULTS:

  • WILD: no data found yet.
  • FARM: not aggressive under high-standard farming conditions 16.
  • LAB: no data found yet.

SPAWNERS:

  • WILD: no data found yet.
  • FARM: not aggressive under high-standard farming conditions 16.
  • LAB: no data found yet.



7  Substrate

Depending on where in the water column the species lives, it differs in interacting with or relying on various substrates for feeding or covering purposes (e.g., plants, rocks and stones, sand and mud, turbidity).

What is the probability of providing the species' substrate and shelter needs in captivity?

It is low for minimal farming conditions, as a) probably all age classes of the species use substrate, but tanks and raceways are devoid of it and b) given stripping in SPAWNERS. It is medium for high-standard farming conditions given a) earthen ponds (which are not replaced by concrete bottom) for JUVENILES to ADULTS, but b) no natural reproduction imaginable for SPAWNERS (which would require spawning substrate in earthen ponds). Our conclusion is based on a low amount of evidence.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS:

  • WILD: found on pebbles with good oxygenation 7↶1.
  • FARM: no data found yet.
  • LAB: no data found yet.

LARVAE and FRY:

  • WILD: Acipenser gueldenstaedtii uses pebbles and stones for concealment 25. Further research needed to determine whether this applies to Acipenser naccarii as well.
  • FARM: for details of holding systems → F1.
  • LAB: no data found yet.

JUVENILES:

  • WILD: BENTHIC 8 on muddy or sandy bottoms 9.
  • FARM: for details of holding systems → F1.
  • LAB: no data found yet.

ADULTS:

  • WILD: → JUVENILES.
  • FARM: for details of holding systems → F1.
  • LAB: no data found yet.

SPAWNERS:

  • WILD: spawn on pebbles in rivers with good oxygenation 7↶1.
  • FARM: manual stripping 13↶26 4 under anaesthesia 4. For details of holding systems → F1.
  • LAB: no data found yet.



8  Stress

Farming involves subjecting the species to diverse procedures (e.g., handling, air exposure, short-term confinement, short-term crowding, transport), sudden parameter changes or repeated disturbances (e.g., husbandry, size grading).

What is the probability of the species not being stressed?

It is low for minimal farming conditions, as the species is likely stressed (water quality, handling, density). It is medium for high-standard farming conditions, as ways to reduce (and avoid?) stress need to be verified for the farming context. Our conclusion is based on a low amount of evidence, as further research is needed.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS:

  • WILD: no data found yet.
  • FARM: no data found yet.
  • LAB: no data found yet.

LARVAE and FRY:

  • WILD: no data found yet.
  • FARM: no data found yet.
  • LAB: no data found yet.

JUVENILES:

  • WILD: in rivers, preference for areas of still water and slower flow 10.
  • FARM: no difference in growth between IND raised in fresh water and water at 11 ppt salinity over 9 weeks 2. Low densities might reduce stress 16. No elevated blood cortisol after netting and handling, possibly due to habituation to human presence 20.
  • LAB: difficult assessment of stress due to large ranges of cortisol levels 20; higher average blood cortisol at 25 °C than 17 °C (within-subject design, sampled after anaesthesia), and no increase in blood cortisol or glucose in IND kept at 17 °C when subjected to stress (crowding for 24 h at 16 kg/m3, long handling; sampled after anaesthesia) 20.

ADULTS:

  • WILD: in rivers, preference for areas of still water and slower flow 10.
  • FARM: low densities might reduce stress 16. Higher blood cortisol after netting and handling (than control), possibly due to isolation from human presence in outdoor tanks 6.
  • LAB: anaesthesia did not reduce cortisol and glucose levels in IND netted and handled, but difficult to interpret results due to large range of blood stress levels 6.

SPAWNERS:

  • WILD: no data found yet.
  • FARM: low densities might reduce stress 16.
  • LAB: no data found yet.



9  Malformations

Deformities that – in contrast to diseases – are commonly irreversible may indicate sub-optimal rearing conditions (e.g., mechanical stress during hatching and rearing, environmental factors unless mentioned in crit. 3, aquatic pollutants, nutritional deficiencies) or a general incompatibility of the species with being farmed.

What is the probability of the species being malformed rarely?

It is unclear for minimal and high-standard farming conditions, given we mainly found unspecific data for one age class. Our conclusion is based on a low amount of evidence, as further research is needed on malformation rates in farms for more age classes.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS:

  • WILD: no data found yet.
  • FARM: hatching rate 66% 3.
  • LAB: no data found yet.

LARVAE and FRY:

  • WILD: no data found yet.
  • FARM: malformation rate at hatching 6.4% 3. Malformations of sensory, circulatory, and digestive systems 15 27, generally lethal, as malformed IND are not found after the stage of FRY 0.
  • LAB: no data found yet.

JUVENILES:

  • WILD: no data found yet.
  • FARM: no data found yet.
  • LAB: no data found yet.

ADULTS:

  • WILD: no data found yet.
  • FARM: no data found yet.
  • LAB: no data found yet.

SPAWNERS:

  • WILD: no data found yet.
  • FARM: no data found yet.
  • LAB: no data found yet.



10  Slaughter

The cornerstone for a humane treatment is that slaughter a) immediately follows stunning (i.e., while the individual is unconscious), b) happens according to a clear and reproducible set of instructions verified under farming conditions, and c) avoids pain, fear, and distress.

What is the probability of the species being slaughtered according to a humane slaughter protocol?

It is low for minimal farming conditions, given the possibility of asphyxia and/or hypothermia (which needs to be verified for Acipenser naccarii). It is high for high-standard farming conditions, given humane stunning followed by slaughter while still insensible and unconscious through (non-penetrating) percussive stunning followed by bleeding, which is verified for the farming context. Our conclusion is based on a low amount of evidence, as further research is needed to confirm the findings.

Likelihoodscore-li
Potentialscore-po
Certaintyscore-ce

EGGS: does not apply.

LARVAE and FRY: does not apply.

JUVENILES:

  • WILD: does not apply.
  • FARM: minimal slaughter method: for the related Acipenser baerii, asphyxia and/or hypothermia by immersion in ice-water slurry 28. Further research needed to determine whether this applies to Acipenser naccarii as well. High-standard slaughter method: percussive stunning through manual spiking, percussive gun 4, or captive bolt pistol 29 performed by experienced staff, followed by bleeding 4.
  • LAB: no data found yet.

ADULTS:

  • WILD: does not apply.
  • FARM: probably → JUVENILES, but sources do not specify the age class 0.
  • LAB: no data found yet.

SPAWNERS:

  • WILD: does not apply.
  • FARM: probably → JUVENILES, but sources do not specify the age class 0.
  • LAB: no data found yet.



Side note: Domestication

Teletchea and Fontaine introduced 5 domestication levels illustrating how far species are from having their life cycle closed in captivity without wild input, how long they have been reared in captivity, and whether breeding programmes are in place.

What is the species’ domestication level?

DOMESTICATION LEVEL 5 30, level 5 being fully domesticated.

While we do not know for sure that Acipenser naccarii reproduction does not entail wild inputs, we know that it is farmed and that hybrids with the related species Acipenser baerii are commonly used in aquaculture. We then consider that Acipenser naccarii should have the same level of domestication as Acipenser baerii 0.




Side note: Forage fish in the feed

450-1,000 milliard wild-caught fishes end up being processed into fish meal and fish oil each year which contributes to overfishing and represents enormous suffering. There is a broad range of feeding types within species reared in captivity.

To what degree may fish meal and fish oil based on forage fish be replaced by non-forage fishery components (e.g., poultry blood meal) or sustainable sources (e.g., soybean cake)?

All age classes:

  • WILD: carnivorous, almost exclusively invertebrates 31.
  • FARM: no replacement of fish meal reported in literature 0. Supplemented linseed oil in diet reduced growth and is not a suitable substitute for fish oil over 1 year of treatment 32.
  • LAB: no data found yet.

*partly = <51%, mostly = 51-99%, completely = 100%




Side note: Commercial relevance

How much is this species farmed annually?

23 t in 2024 33.




Glossary

ADULTS = mature individuals
ANADROMOUS = migrating from the sea into fresh water to spawn
BENTHIC = living at the bottom of a body of water, able to rest on the floor
DOMESTICATION LEVEL 5 = selective breeding programmes are used focusing on specific goals 30
DPH = days post hatching
EGGS = reproductive product containing - if fertilised - a ripening embryo until hatching
FARM = setting in farming environment or under conditions simulating farming environment in terms of size of facility or number of individuals
FRY = larvae from external feeding on
IND = individuals
JUVENILES = fully developed but immature individuals
LAB = setting in laboratory environment
LARVAE = hatching to mouth opening
POTAMODROMOUS = migrating within fresh water
SEMI-ANADROMOUS = migrating from downstream or the estuary into fresh water upstream to spawn
SPAWNERS = adults during the spawning season; in farms: adults that are kept as broodstock
WILD = setting in the wild



Bibliography

0 Own conclusion
1 Conservation and Breeding of Italian Cobice Endemic Sturgeon (COBICE). 2007. Report 04NAT/IT/000126. Italy: Ente Parco Regionale Veneto del Delta del Po.
2 McKenzie, D J, E Cataldi, P Romano, S F Owen, E W Taylor, and P Bronzi. 2001. Effects of acclimation to brackish water on the growth, respiratory metabolism, and swimming performance of young-of-the-year Adriatic sturgeon ( Acipenser naccarii ). Canadian Journal of Fisheries and Aquatic Sciences 58: 1104–1112. https://doi.org/10.1139/f01-059.
3 Cabrera-Castro, R., C. Zabala, M.C. Soriguer, A. Domezain, and J.A. Hernando. 2018. Morphological development in the first life phase of Adriatic sturgeon Acipenser  naccarii under controlled conditions. Journal of Fish Biology 92: 1956–1974.
4 Saraiva, João L. 2018. Personal communication.
5 Furné, M., M. García-Gallego, M.C. Hidalgo, A.E. Morales, A. Domezain, J. Domezain, and A. Sanz. 2009. Oxidative stress parameters during starvation and refeeding periods in Adriatic sturgeon ( Acipenser naccarii ) and rainbow trout ( Oncorhynchus mykiss ). Aquaculture Nutrition 15: 587–595. https://doi.org/10.1111/j.1365-2095.2008.00626.x.
6 Di Marco, P., D.J. McKenzie, A. Mandich, P. Bronzi, E. Cataldi, and S. Cataudella. 1999. Influence of sampling conditions on blood chemistry values of Adriatic sturgeon Acipenser naccarii (Bonaparte, 1836). Journal of Applied Ichthyology 15: 73–77. https://doi.org/10.1111/j.1439-0426.1999.tb00210.x.
7 Holčík, Juraj. 1989. The Freshwater Fishes of Europe: General introduction to fishes ; Acipenseriformes. AULA - Verlag.
8 Rossi, R, G Grandi, R Trisolini, P Franzoi, A Carrieri, BS Dezfuli, and E Vecchietti. 1991. Osservazioni sulla biologia e la pesca dello storione cobice Acipenser naccarii Bonaparte nella parte terminale del fiume Po. Atti della Società italiana di scienze naturali e del museo civico di storia naturale di Milano 132: 121–142.
9 Bronzi, Paolo, Paul Vecsei, and Giovanni Arlati. 2005. Threatened fishes of the world: Acipenser naccarii Bonaparte, 1836 (Acipenseridae). Environmental Biology of Fishes 72: 66–66. https://doi.org/10.1007/s10641-004-1036-4.
10 Labria, F., V. Orioli, S. Trasforini, C. Puzzi, P. Tirozzi, L. Bani, and O. Dondina. 2025. Unlocking the Secrets of Sturgeon Ecology: Lessons From Conservation Efforts in Northern Italy. Aquatic Conservation: Marine and Freshwater Ecosystems 35.
11 Pagani, S., G. Castaldelli, L. Congiu, P. Bronzi, R. Rossi, and M. Lanzoni. 2026. The Record of the Last Sturgeons Caught in the Po River (North Italy) Tells a Cautionary Tale of Reasons of Their Silent Disappearance. River Research and Applications 42: 277–283.
12 Arlati, G. 1996. Ripopolamento con lo storione nelle acque pubbliche. In , 11.
13 Arlati, G., P. Bronzi, L. Colombo, and G. Giovannini. 1988. Induced breeding of the Italian Sturgeon (Acipenser naccarii) reared in captivity. Rivista Italiana di Acquacultura 23: 94–96.
14 Arlati, G., A. Grassi, and A. Granata. 1999. Restocking Acipenser Naccarii in the Lombardy Region. Journal of Applied Ichthyology 15: 298–298. https://doi.org/10.1111/j.1439-0426.1999.tb00288.x.
15 Boglione, C., P. Bronzi, E. Cataldi, S. Serra, F. Gagliardi, and S. Cataudella. 1999. Aspects of early development in the Adriatic sturgeon Acipenser naccarii. Journal of Applied Ichthyology 15: 207–213. https://doi.org/10.1111/j.1439-0426.1999.tb00236.x.
16 Anonymous farmer. 2026. Personal communication.
17 Anonymous farmers. 2018. Personal communication.
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